My Research Interests and History: I received my Ph.D. in 1993 with Richard Root at Cornell University, and performed my postdoctoral studies with David Tilman at the University of Minnesota and Steve Hubbell at Princeton University.  I joined the University of Pittsburgh Department of Biological Sciences in 1994 and retired in 2023. I am currently an Adjunct Professor at the University of Nevada, Reno.


The research in my lab continues to focus on four major areas: